500
to 1900
Migration · Arabian
Iranian Migration to Eastern Arabia
Ajam and Huwala merchants and settlers introduced Iranian-related ancestry along the Gulf, with diffusion inland through marriage and patronage ties.
Source confidence: high
500
to 1750
Migration · Indian
Jewish migrations to Cochin and Bombay
Small communities retained Levantine ancestry with admixture from local populations. Genetic signatures persist due to partial endogamy.
Source confidence: medium
525
to 570
Civilization · Arabian
Aksumite Rule in Yemen
Aksumite garrisons and merchants settled locally, contributing Horn of Africa ancestry to Yemeni coastal and adjacent highland populations.
Source confidence: high
525
to 570
Migration · Arabian
Aksumite Migration into Yemen
Military settlers and administrators from the Horn of Africa contributed East African ancestry, especially along Yemen’s coastal-lowland belt.
Source confidence: medium
533
to 698
Civilization · North African
Byzantine Exarchate of Africa
Reintegration into the Eastern Mediterranean sphere restored flows of Greek, Levantine, and Anatolian personnel. Admixture layered onto existing Romanized Amazigh populations in port cities.
Source confidence: medium
534
to 900
Civilization · Sardinian
Byzantine Sardinia (Theme of Sardinia)
Small Eastern Mediterranean (Greek/Anatolian) inputs via officials, soldiers, and clergy. Admixture concentrated in towns, with rural interiors remaining relatively isolated.
Source confidence: medium
534
to 900
Migration · Sardinian
Byzantine officials, soldiers, and clergy arriving in Sardinia
Added modest Eastern Mediterranean input concentrated in administrative and ecclesiastical hubs. Rural interiors remained comparatively isolated.
Source confidence: medium
550
to 800
Civilization · Indonesian Khmer Thai Myanma
Chenla
Consolidation of inland Austroasiatic-speaking groups around the Mekong facilitated gene flow among Khmer-related populations and with neighboring highlanders. Elite integration likely spread Khmer-affiliated lineages.
Source confidence: medium
550
to 1000
Civilization · Indonesian Khmer Thai Myanma
Dvaravati (Mon polities)
Mon-speaking communities established urban centers, contributing Austroasiatic ancestry to central Thai populations. Subsequent incorporation by Tai states produced sustained admixture with incoming Tai groups.
Source confidence: medium
570
to 630
Civilization · Arabian
Sasanian Yemen
Iranian administrators and soldiers established colonies in ports and oases, introducing Persian Gulf-related lineages that partly assimilated via marriage.
Source confidence: medium
581
to 618
Civilization · Chinese
Sui Dynasty
Political unification and canal connectivity increased mobility between northern and southern populations. Population movements promoted mixing across previously separated gene pools.
Source confidence: medium
600
to 900
Migration · Chinese
Tang-era Silk Road Immigrants
Central and West Asian lineages entered corridor and urban populations, leaving minor but detectable western Eurasian signals. Many descendants assimilated into Han and Hui communities.
Source confidence: medium
600
to 1500
Migration · Indian
Arab and Persian merchant settlement on Indian coasts
Male-mediated West Asian gene flow entered coastal Muslim communities (e.g., Mappila, Bohra, Khoja). Admixture signals are strongest in port cities.
Source confidence: high
611
to 628
Civilization · Levantine
Sasanian Empire (occupation of the Levant)
Brief Sasanian control introduced garrisons and administrators from Iran and the Caucasus. Lasting genetic impact was likely small and localized around occupied cities.
Source confidence: low
618
to 907
Civilization · Chinese
Tang Dynasty
Cosmopolitan Silk Road cities received Turkic, Sogdian, and Persian immigrants who admixed into urban and corridor populations. Expansion and garrisons also spread Han lineages and fostered mixing in the northwest and southwest.
Source confidence: high
630
to 800
Migration · Levantine
Arab tribal migrations during Islamic conquests and settlement
Conquest-era tribal garrisons and their families settled across the Levant, intermarrying with local populations. This established enduring Arabian-associated ancestry components in many communities.
Source confidence: high
630
to 850
Migration · Chinese
Tibetan Imperial Expansion on the Plateau
Consolidated Tibetan-related ancestry and mixed with Tang settlers and Qiangic groups. This shaped modern Tibetan, Amdo, and adjacent highland gene pools.
Source confidence: medium
632
to 750
Civilization · Arabian
Rashidun and Umayyad Caliphates
Mass tribal mobilization and new settlements spread central Arabian lineages while drawing in people from the Levant and Mesopotamia through military, administrative, and familial ties.
Source confidence: high
632
to 900
Migration · Arabian
Early Islamic Iraqi and Levantine Influx
Administrators, scholars, and soldiers moved into Arabian centers during early Islam, adding Levantine and Mesopotamian ancestry particularly in the Hejaz and Gulf.
Source confidence: medium
634
to 661
Civilization · Levantine
Rashidun Caliphate
Conquest and organized tribal settlement from Arabia established new Arab communities and accelerated Arabicization. Admixture occurred through intermarriage and conversion within towns and rural districts.
Source confidence: high
639
to 750
Civilization · North African
Rashidun and Umayyad Caliphates in North Africa
Arab garrisons and settlers established new towns and tribal quarters, introducing Southwest Asian ancestry. Admixture with Amazigh and Coptic communities accompanied the spread of Arabic and Islam.
Source confidence: high
639
to 800
Migration · North African
Arab Conquest and Settlement of Egypt and the Maghreb
Introduced Southwest Asian ancestry with male-biased inputs in garrison towns. Admixture with Amazigh and Coptic populations underpinned the emergence of Arabic-speaking communities.
Source confidence: high
639
to 969
Civilization · Egyptian
Early Islamic Caliphates in Egypt (Rashidun–Abbasid)
Arab tribal settlers and administrators from Arabia and the Levant founded Fustat and rural garrisons. Admixture with local Copts and ongoing Levantine links introduced Arabian and broader Eastern Mediterranean ancestry.
Source confidence: high
639
to 1000
Migration · Egyptian
Arab Conquest and Settlement of Egypt
Migration of Arab tribes, garrisons, and administrators introduced Arabian and Levantine ancestry, with male-biased inputs into patrilines. Assimilation with Coptic populations spread these components broadly, strongest in northern and urban locales.
Source confidence: high
650
to 1300
Civilization · Indonesian Khmer Thai Myanma
Srivijaya
Maritime entrepôts facilitated sustained South Asian, Arab, and Chinese gene flow into Malay-Austronesian coastal groups. Mobility within the thalassocracy mixed Sumatran, Javanese, and Bornean populations.
Source confidence: high
661
to 750
Civilization · Levantine
Umayyad Caliphate
The Damascus-centered state consolidated earlier Arab settlement and drew people from Arabia, Iraq, and beyond into the Levant. Court, army, and trade networks fostered gene flow among Arab, local Levantine, and other West Asian groups.
Source confidence: high
700
to 1900
Migration · Levantine
African slave trade into Levantine households and markets
Trans-Saharan and Red Sea routes brought Africans to Levantine ports and cities; many assimilated over generations. This contributed low-to-moderate Sub-Saharan and Nilotic ancestry in some coastal and urban groups.
Source confidence: medium
700
to 1900
Migration · North African
Trans-Saharan Slave Trade to North Africa
Sustained movements introduced substantial sub-Saharan ancestry, particularly in oasis zones and urban centers; maternal lineages show pronounced signals from this period.
Source confidence: high
700
to 1900
Migration · Arabian
Indian Ocean Slave Trade into Arabia
Enslaved Africans and their descendants integrated into coastal and urban societies, leaving notable African ancestry, often via maternal lines, in many littoral populations.
Source confidence: high
700
to 2000
Migration · Arabian
Hajj Pilgrim Settlement in Hejaz
A steady stream of pilgrims remained in the holy cities, creating a highly mixed urban gene pool that reflects broad Afro-Eurasian origins.
Source confidence: high
700
to 1600
Migration · Chinese
Hui Muslim Formation via Intermarriage and Conversion
Admixture of foreign Muslim lineages with Han and other local groups created Hui populations with predominantly East Asian ancestry and minor western Eurasian components. Community endogamy maintained distinct identity.
Source confidence: high
700
to 1100
Migration · Indian
Parsi (Zoroastrian) migration to Gujarat
Established endogamous communities retaining Iranian-related ancestry with subsequent admixture from local Gujaratis. Genetic distinctiveness persists alongside South Asian components.
Source confidence: high
700
to 1600
Migration · Indonesian Khmer Thai Myanma
Tai migrations into mainland Southeast Asia
Tai-speaking groups admixed with Mon-Khmer and other local populations, forming the core of Thai and Shan gene pools. Elite-state formation amplified Tai ancestry in lowland valleys.
Source confidence: high
700
to 1200
Migration · Indonesian Khmer Thai Myanma
Bamar expansion into the Irrawaddy plains
Consolidated Tibeto-Burman ancestry with local Pyu and Mon groups in central Myanmar. Established demographic cores that continued to absorb neighboring populations.
Source confidence: high
700
to 1880
Migration · Egyptian
Trans-Saharan Slave Trade to Egypt
Centuries of enslaved and manumitted individuals from the Sahel and Central Africa contributed sub-Saharan ancestry to both urban and rural Egyptian gene pools. Gene flow was often female-biased in domestic contexts and male-biased in military cohorts.
Source confidence: high
738
to 1253
Civilization · Chinese
Nanzhao-Dali Kingdoms
Integration of Bai, Yi, and Tai groups with migrants from Tang and Song produced distinctive southwest Chinese admixture. Subsequent Mongol conquest added steppe components to Yunnanese populations.
Source confidence: medium
750
to 969
Civilization · Levantine
Abbasid Caliphate (Syrian provinces)
Though the imperial center moved east, garrisons, officials, and merchants continued to circulate between Iraq, Egypt, and the Levant. These links maintained West Asian and Nile-to-Levant genetic exchanges.
Source confidence: medium
750
to 1258
Civilization · Arabian
Abbasid Caliphate Influence in Arabia
Pilgrimage, scholarship, and Gulf commerce increased inflow from Iraq, Iran, and the wider Islamic world; enslaved and merchant populations added diversity to coastal and holy cities.
Source confidence: high
750
to 1174
Civilization · Indian
Pala Empire
Buddhist scholastic and trade networks linked Bengal to the Himalayas and Southeast Asia, fostering movement of monks and artisans. Regional urbanization encouraged admixture within eastern India.
Source confidence: medium
752
to 1045
Civilization · Indonesian Khmer Thai Myanma
Medang (Mataram) Kingdom
State formation and shifts of population centers within Java promoted mixing among Javanese communities and with Balinese. Limited South Asian inputs arrived via religious and trade ties.
Source confidence: medium
800
to 950
Migration · Northwestern European
Norse settlement in the Lower Seine (formation of Normandy)
Viking groups settled and assimilated, contributing Scandinavian male-line ancestry while adopting local customs and language. The result was a mixed Norse–Frankish population base.
Source confidence: high
800
to 1506
Migration · South East African
Swahili–Arab coastal settlement in Mozambique
Maritime settlers and merchants intermarried with coastal Bantu communities, introducing West Asian ancestry and Islamic cultural affiliations.
Source confidence: high
800
to 909
Civilization · North African
Aghlabid Emirate of Ifriqiya
Further Arabization of coastal Tunisia and expansion into Sicily facilitated bidirectional movement of people across the central Mediterranean. Urban centers saw continued mixing of Amazigh, Arab, and Mediterranean groups.
Source confidence: medium
800
to 1800
Migration · African Hunter-Gatherer
Nilotic Pastoralist Migrations into East Africa
Nilotic groups expanded southward, interacting with and sometimes absorbing local foragers on the savanna–forest margins. Resulting gene flow is detectable in adjacent agro-pastoral populations and some forager neighbors.
Source confidence: medium
800
to 1800
Migration · Indian
African (Siddi/Habshi) arrivals via the Indian Ocean
Brought sub-Saharan African ancestry to coastal and Deccan regions through slave trade and military service. Enduring communities show substantial African-derived components.
Source confidence: high
802
to 1431
Civilization · Indonesian Khmer Thai Myanma
Khmer Empire (Angkor)
State projects and warfare drove internal relocations and assimilation of Mon, Cham, and Tai groups into Khmer cores. These movements increased regional genetic connectivity across the Mekong basin.
Source confidence: high
849
to 1297
Civilization · Indonesian Khmer Thai Myanma
Pagan (Bagan) Kingdom
Bamar expansion integrated Pyu and Mon populations, consolidating Tibeto-Burman and Austroasiatic ancestries in central Myanmar. Monastic networks and urban growth fostered continued mixing along the Irrawaddy.
Source confidence: high
850
to 1279
Civilization · Indian
Chola Empire
Maritime power expanded port cities and guild networks, attracting foreign merchants and migrants. Coastal communities show signals of admixture facilitated by long-distance trade.
Source confidence: medium
900
to 1300
Migration · Northwestern European
Ashkenazi Jewish settlement in the Rhineland
Founding communities established enduring Jewish populations with characteristic founder effects and endogamy. These groups later dispersed eastward but left a detectable regional legacy.
Source confidence: high
900
to 1323
Civilization · Sardinian
Giudicati of Sardinia (Judicates)
Local rule sustained endogamy and continuity of earlier lineages, preserving regional substructure. External inputs occurred mainly through elite marriages and merchants.
Source confidence: medium
907
to 1125
Civilization · Chinese
Liao Dynasty (Khitan)
Khitan elites and settlers introduced steppe ancestry into northeast China and adjacent plains. Dual administration and intermarriage produced localized admixture with Han and other groups.
Source confidence: medium
907
to 1125
Migration · Chinese
Khitan Migrations under the Liao
Introduced Khitan-related steppe ancestry into northeast and northern frontier populations. Admixture occurred via garrisons and mixed households.
Source confidence: medium
909
to 1171
Civilization · North African
Fatimid Caliphate (Ifriqiya and Egypt)
Kutama Berber armies and administrators moved east to Egypt, increasing Maghreb–Egypt gene flow. Slave-soldiers and servants from sub-Saharan Africa and Eurasia added diversity to urban populations.
Source confidence: high
911
to 1204
Civilization · Northwestern European
Duchy of Normandy (Norse-descended polity)
Scandinavian settlers became a dominant elite that integrated with local populations. Male-line Scandinavian input persisted while overall admixture aligned Normandy with northern France.
Source confidence: medium
960
to 1279
Civilization · Chinese
Song Dynasty
A demographic center shift to the south intensified admixture between Han migrants and indigenous Tai-Kadai, Hmong-Mien, and Austronesian-related populations. Maritime and inland trade sustained limited additional inputs.
Source confidence: high
962
to 1806
Civilization · Northwestern European
Holy Roman Empire
Imperial networks of cities, monasteries, and armies promoted mobility of artisans, merchants, and soldiers. This facilitated gene flow among regional German-speaking and Romance-speaking communities.
Source confidence: high
967
to 1916
Civilization · Arabian
Sharifate of Mecca
Continuous Hajj-centered settlement by pilgrims and scholars from Africa and Asia created a cosmopolitan gene pool in Hejazi urban centers.
Source confidence: high
969
to 1099
Civilization · Levantine
Fatimid Caliphate
Administration and garrisons from Egypt and the Maghreb, along with slave-soldier imports from Africa and the Caucasus, added small North African and African-associated components to coastal and urban populations.
Source confidence: medium
969
to 1171
Civilization · Egyptian
Fatimid Caliphate in Egypt
Berber and other Maghrebi contingents accompanied the Fatimids, while slave-soldier and domestic networks brought sub-Saharan and Caucasus individuals to Cairo. These streams modestly diversified urban gene pools, especially in Fustat/Cairo.
Source confidence: medium
987
to 1589
Civilization · Northwestern European
Kingdom of France (Capetian–Valois)
State consolidation and growth of towns drove internal migration and mixing across northern France. Integration of Burgundy and Champagne linked eastern French populations to the frontier zone.
Source confidence: medium
1000
to 1506
Civilization · South East African
Kilwa Sultanate and Sofala Port
Coastal communities show admixture between Bantu-speaking groups and migrants of Arab and Persian origin, with male-mediated gene flow prominent. Islamic networks fostered exogamy and long-distance kin ties along the coast.
Source confidence: high
1000
to 1800
Migration · South East African
Madagascar–Mozambique Channel interactions
Cross-channel voyages and raiding produced modest Malagasy/Austronesian gene flow into coastal Mozambique and reciprocal African ancestry in western Madagascar.
Source confidence: medium
1000
to 1900
Migration · North African
Tuareg Saharan Pastoral Movements
Mobile Amazigh groups redistributed Saharan ancestry and mediated additional sub-Saharan gene flow into northern fringes via caravan networks and alliances.
Source confidence: medium
1000
to 2000
Migration · Arabian
South Asian Merchant and Artisan Migration
Maritime commerce brought South Asian lineages to Arabian port towns, with assimilation into coastal families and trading diasporas.
Source confidence: medium
1000
to 1900
Migration · Chinese
Hakka Intra-Southern Migrations
Serial movements created founder effects and substructure while mixing with local southern groups. Resulting Hakka populations show distinctive yet regionally admixed profiles.
Source confidence: medium
1000
to 1800
Civilization · African Hunter-Gatherer
Swahili City-States
Coastal trade and slaving drew individuals from inland forests to the coast, occasionally relocating foragers and mediating admixture with coastal populations. Most gene flow into foragers was indirect via inland partners tied to caravans.
Source confidence: medium
1000
to 1900
Migration · African Hunter-Gatherer
Indian Ocean Slave Trade and Inland Caravans
Capture and sale of inland peoples, including some forest foragers, moved individuals to coastal and overseas populations. This reduced forager population sizes and introduced forager lineages into coastal communities.
Source confidence: medium
1000
to 1870
Civilization · Papuan
Motu Hiri Trade Complex
Seasonal pottery and sago exchange voyages fostered intermarriage between Motu (Austronesian) and neighboring Papuan groups. This increased Austronesian ancestry on the south coast while maintaining localized structure.
Source confidence: medium
1000
to 2000
Civilization · Papuan
Polynesian Outlier Communities (Nuguria, Nukumanu, Takuu)
Polynesian settlers established endogamous communities on PNG atolls, preserving Polynesian ancestry. Occasional intermarriage with nearby Melanesian groups produced localized Polynesian–Papuan admixture.
Source confidence: high
1000
to 1500
Migration · Papuan
Polynesian Outlier Colonizations of PNG Atolls
Voyaging parties established Polynesian communities on remote atolls, maintaining endogamy with episodic marriage exchanges with nearby Melanesians. Resulting gene pools retain strong Polynesian signals with localized admixture.
Source confidence: high
1017
to 1070
Migration · Indonesian Khmer Thai Myanma
Chola expeditions and merchant-soldier settlements
Short-term occupations and fortified trade posts seeded small South Indian communities that intermarried locally, adding limited South Asian signals to coastal gene pools.
Source confidence: medium
1038
to 1227
Civilization · Chinese
Western Xia (Tangut)
Tangut rule integrated Tibeto-Burman, Han, and steppe communities in the Hexi Corridor. This produced enduring admixture signals in northwestern populations.
Source confidence: medium
1040
to 1147
Civilization · North African
Almoravid Dynasty
Saharan Amazigh groups moved northward, knitting together desert and coastal populations. Ties to al-Andalus promoted additional Mediterranean admixture in Maghrebi cities.
Source confidence: medium
1050
to 1200
Migration · North African
Banu Hilal and Banu Sulaym Migrations
Nomadic Arab tribes dispersed across rural Maghreb, accelerating linguistic Arabization and adding Arabian patrilines to pastoral and agro-pastoral populations.
Source confidence: medium
1050
to 1326
Civilization · Sardinian
Pisan Influence and Control
Tuscan colonists and miners added Ligurian-Tuscan ancestry to coastal and mining towns. Impact was localized, with limited penetration into the interior.
Source confidence: medium
1070
to 1400
Migration · Levantine
Turkic and Turkmen migrations into Syria and the Levant
Seljuk and Turkmen groups established emirates and pastoral camps, later integrating into local societies. Their presence introduced Central Asian/Turkic-associated Y-chromosome and autosomal components.
Source confidence: medium
1075
to 1220
Civilization · South East African
Kingdom of Mapungubwe
Aggregation of agropastoral populations around a hierarchical center increased connectivity and gene flow across the Limpopo corridor. Contacts with coastal trade routes linked inland groups to broader exchange networks.
Source confidence: medium
1099
to 1291
Civilization · Levantine
Crusader States (Latin East)
Frankish settlers and military orders established enclaves that intermarried with local Christians and Muslims. Genetic inputs from Western Europe persisted at low frequencies in some coastal and highland communities.
Source confidence: medium
1099
to 1291
Migration · Levantine
Crusader/Frankish settlement
Tens of thousands of settlers formed colonial enclaves and intermarried with local Christians and Muslims. Western European genetic inputs entered some coastal and inland lineages at low frequencies.
Source confidence: medium
1100
to 1326
Civilization · Sardinian
Genoese Influence and Colonies
Ligurian merchant families and settlers contributed gene flow to northern coastal communities. Effects are most visible in port towns and mercantile lineages.
Source confidence: medium
1100
to 1300
Migration · Sardinian
Pisan-Tuscan colonists and miners
Contributed Tuscan/Ligurian ancestry to coastal and mining populations. Impact was localized to towns and industrial enclaves.
Source confidence: medium
1100
to 1916
Civilization · African Hunter-Gatherer
Kingdom of Rwanda
Twa foragers were incorporated at the margins of a stratified kingdom, producing limited but measurable admixture with neighboring agricultural populations. Social constraints reduced but did not eliminate gene flow.
Source confidence: high
1115
to 1234
Civilization · Chinese
Jin Dynasty (Jurchen)
Jurchen conquest brought Manchurian-related ancestry into northern populations and implemented relocations that reshaped urban and rural gene pools. Mixed households and garrisons reinforced admixture.
Source confidence: high
1121
to 1269
Civilization · North African
Almohad Caliphate
Imperial consolidation spurred movement of tribes, administrators, and artisans across the Maghreb. This increased homogenization among Amazigh- and Arab-speaking groups with continued Andalusi inputs.
Source confidence: high
1125
to 1234
Migration · Chinese
Jurchen Expansion and Population Transfers
Conquest and relocations increased Manchurian-related admixture in northern China and displaced many Han southward. Urban centers show signals of mixed Jurchen-Han ancestry.
Source confidence: high
1170
to 1900
Migration · Levantine
Kurdish resettlement under Ayyubids and Ottomans
Military colonization and tribal movements placed Kurdish communities across northern Levant. Over centuries, intermarriage diffused Upper Mesopotamian genetic signatures into nearby populations.
Source confidence: medium
1171
to 1250
Civilization · Levantine
Ayyubid Sultanate
Kurdish-led dynasties moved Kurdish, Turkic, and other military households into Syrian cities and countryside. Their retinues and alliances introduced additional Upper Mesopotamian and steppe-associated lineages.
Source confidence: medium
1171
to 1250
Civilization · Egyptian
Ayyubid Sultanate in Egypt
Kurdish-led elites and Turkic auxiliaries settled and intermarried within urban society, while recruitment of slaves from the Caucasus and Africa continued. Inputs were male-biased and concentrated in cities and military households.
Source confidence: medium
1200
to 1450
Civilization · South East African
Kingdom of Zimbabwe (Great Zimbabwe)
Consolidation of Shona-speaking populations promoted admixture among plateau communities. Trade with the coast facilitated gene flow between inland and coastal populations.
Source confidence: high
1200
to 1517
Migration · North African
Mamluk Slave-Soldier Imports to Egypt
Introduced Caucasus and steppe ancestries into Egypt’s urban elite, with diffusion into broader populations through descendants and social integration.
Source confidence: medium
1200
to 2000
Migration · Arabian
Hadrami Diaspora and Return Flows
Overseas Hadrami communities intermarried locally and sent spouses and descendants back, introducing Southeast Asian and East African lineages into Hadramawt and Yemen’s coasts.
Source confidence: medium
1200
to 1900
Civilization · African Hunter-Gatherer
Kingdom of Buganda
Expansion toward forest margins brought Batwa and other forager-descended groups into tributary systems, enabling sporadic admixture with Bantu-speaking farmers. Gene flow was strongest along lake and forest-edge corridors.
Source confidence: medium
1200
to 1900
Migration · Indonesian Khmer Thai Myanma
Hadhrami Arab diaspora to the Indonesian archipelago
Male-mediated gene flow introduced Arabian patrilines and contributed to Islamic scholarly and elite networks, with admixture concentrated in trading cities and among coastal elites.
Source confidence: medium
1200
to 2000
Civilization · Papuan
Massim/Trobriand Chiefdoms and the Kula Exchange
Long-distance ceremonial exchange networks promoted maritime mobility and affinal ties across islands. These processes maintained island-specific lineages while enabling periodic gene flow between Austronesian and Papuan communities.
Source confidence: high
1206
to 1526
Civilization · Indian
Delhi Sultanate
Turkic, Afghan, and Persian migrants introduced West/Central Asian ancestry, especially in urban Muslim populations. Population transfers and slavery redistributed people across North India.
Source confidence: high
1206
to 1526
Migration · Indian
Turkic and Afghan influx during the Delhi Sultanate
Soldiers, administrators, and artisans added West/Central Asian ancestry to northern urban populations, often male-biased. Slave-taking and resettlement redistributed people regionally.
Source confidence: high
1211
to 1368
Migration · Chinese
Mongol Resettlements and Semu Migration
Movements of Mongol, Turkic, and Southwest/West Asian groups introduced diverse ancestries, notably in northern cities and Yunnanese populations. Admixture persisted through subsequent dynasties.
Source confidence: high
1221
to 1327
Migration · Indian
Mongol/Chagatai entrants into North India
Small contingents settled as auxiliaries after defections, contributing limited Central Asian ancestry. Overall genetic impact was minor and localized.
Source confidence: low
1228
to 1826
Civilization · Indian
Ahom Kingdom
Tai-Ahom settlers assimilated and intermarried with local Tibeto-Burman and Austroasiatic groups. Resulted in persistent East/Southeast Asian-related ancestry in Assam.
Source confidence: high
1229
to 1454
Civilization · Arabian
Rasulid Dynasty of Yemen
Active Indian Ocean trade under Rasulid rule reinforced South Asian and East African genetic inputs in Yemeni ports and disseminated them into the interior.
Source confidence: medium
1238
to 1438
Civilization · Indonesian Khmer Thai Myanma
Sukhothai Kingdom
Tai-speaking elites established rule over Mon/Khmer substrates, producing admixture that helped form the early Thai genetic profile. Population movement between valleys increased Tai-Mon-Khmer blending.
Source confidence: high
1250
to 1517
Civilization · Levantine
Mamluk Sultanate
Continuous importation of Kipchak and Circassian slaves formed military elites who settled and married locally. This period increased Caucasus and steppe-derived ancestry in urban Levantine populations.
Source confidence: high
1250
to 1517
Migration · Levantine
Mamluk-era Caucasian slave-soldier influx
Large cohorts of Circassian and Kipchak males were brought as slaves, manumitted, and often established families. This added measurable Caucasus-steppe ancestry to urban Levantine populations.
Source confidence: high
1250
to 1517
Civilization · North African
Mamluk Sultanate of Egypt
Recruitment of Caucasian and steppe slave-soldiers introduced new ancestries into Egyptian elites and urban populations. Continued import of African slaves added to genetic diversity, especially maternally.
Source confidence: high
1250
to 1517
Migration · Egyptian
Mamluk Military Slave Imports
Successive cohorts of Caucasus and steppe youths were integrated into military households and urban society, yielding a strong male-line signal in elite and artisan families. Admixture diffused into broader city populations via marriage and patronage networks.
Source confidence: high
1271
to 1368
Civilization · Chinese
Yuan Dynasty (Mongol)
Mongol rule relocated soldiers, artisans, and administrators from Inner Asia and West/Southwest Asia, introducing steppe and minor western Eurasian lineages. Admixture was strongest in northern cities and Yunnan.
Source confidence: high
1273
to 1648
Civilization · Northwestern European
Habsburg Further Austria and Upper Rhine Dominions
Habsburg rule connected Upper Rhine populations with Austrian and Swiss-Alemannic elites and settlers. Cross-border marriages and garrisons modestly increased trans-Alpine genetic ties.
Source confidence: medium
1293
to 1527
Civilization · Indonesian Khmer Thai Myanma
Majapahit Empire
Inter-island administration and commerce drove mobility and intermarriage among Javanese, Balinese, Malay, and eastern Indonesian groups, increasing gene flow along maritime corridors.
Source confidence: high
1300
to 1900
Civilization · African Hunter-Gatherer
Tswana Polities
Tswana chiefdoms incorporated neighboring San through clientage and warfare, leading to sustained Bantu–Kalahari forager admixture. This reshaped gene pools on the Kalahari periphery.
Source confidence: medium
1300
to 1900
Civilization · Papuan
Siassi Maritime Trading Network
Regional traders bridged Papuan and Austronesian populations across the Vitiaz Strait, facilitating movement of people alongside goods. Resulting admixture is detectable in coastal and island communities.
Source confidence: medium
1323
to 1479
Civilization · Sardinian
Crown of Aragon (Kingdom of Sardinia, Aragonese period)
Catalan and Valencian colonization introduced Iberian ancestry, notably in Alghero where Catalan culture persisted. Integration with locals produced urban admixture.
Source confidence: high
1336
to 1646
Civilization · Indian
Vijayanagara Empire
Urban centers and military frontiers drew migrants from across southern India, promoting admixture among Kannada-, Telugu-, and Tamil-speaking groups. Also integrated mercenaries of diverse origins.
Source confidence: high
1347
to 1686
Civilization · Indian
Bahmani and Deccan Sultanates
Influx of Persianate, Turkic, Arab, and African (Habshi/Siddi) soldiers and administrators contributed West Asian and African lineages to Deccan urban and elite strata. Intermarriage and patronage broadened admixture.
Source confidence: high
1351
to 1767
Civilization · Indonesian Khmer Thai Myanma
Ayutthaya Kingdom
Warfare and captive resettlement brought Mon, Khmer, Lao, and Shan groups into Siamese cores, raising heterogeneity. Cosmopolitan ports attracted Chinese, Indian, and Persian migrants who intermarried locally.
Source confidence: high
1354
to 1500
Migration · Sardinian
Catalan-Aragonese colonization (repopulation of Alghero and urban centers)
Catalan settlers created an Iberian enclave at Alghero with lasting genetic and cultural signatures. Smaller Iberian inputs spread to other towns via garrisons and officials.
Source confidence: high
1368
to 1644
Civilization · Chinese
Ming Dynasty
State-sponsored garrisons and agricultural colonies expanded Han ancestry into the southwest and northwest, mixing with Yi, Miao, Zhuang, and Hui communities. Continued commerce sustained localized gene flow from Central and Southeast Asia.
Source confidence: high
1380
to 1650
Migration · Chinese
Ming Tuntian Colonization of the Southwest
Soldier-farmer colonies and migrant households expanded Han ancestry and promoted admixture with Yi, Miao, and Zhuang populations. This reshaped genetic structure across the southwest.
Source confidence: high
1391
to 1614
Migration · North African
Andalusi, Sephardi, and Morisco Dispersals to the Maghreb and Egypt
Refugee and merchant communities added Iberian and mixed Andalusi ancestries to North African cities, expanding Jewish Sephardi and Muslim Morisco-descended lineages.
Source confidence: high
1400
to 1800
Migration · Sardinian
Corsican migration to Gallura and northern Sardinia
Introduced Corsican/Ligurian-related lineages into northern Sardinia, influencing dialects and surname distributions. Gene flow was strongest along the Strait of Bonifacio.
Source confidence: high
1400
to 1888
Civilization · African Hunter-Gatherer
Kingdom of Kongo
Incorporation of forest forager communities through clientage and slavery led to Bantu–hunter-gatherer admixture in western Central Africa. Gene flow was often sex-biased and concentrated along riverine trade corridors.
Source confidence: medium
1400
to 1900
Migration · Indonesian Khmer Thai Myanma
Overseas Chinese migration to Thailand, Myanmar, and Indonesia
Introduced substantial East Asian ancestry into urban and trading communities, forming Sino-Thai, Peranakan, and Chinese-Burmese groups through sustained intermarriage.
Source confidence: high
1450
to 1760
Civilization · South East African
Mutapa State (Mwene Mutapa)
Expansion into the Zambezi valley integrated diverse groups, with intermarriage among Shona elites, valley communities, and coastal/Luso-African families. Movement of specialists and caravans broadened regional admixture.
Source confidence: high
1479
to 1718
Civilization · Sardinian
Spanish Habsburg Rule
Moderate Iberian inflow via officials and soldiers added to earlier Catalan inputs. Genetic effects were concentrated in cities and fortified sites.
Source confidence: medium
1480
to 1779
Civilization · South East African
Maravi Confederacy
Expansion and fragmentation of Maravi-linked groups (e.g., Chewa, Nyanja) increased gene flow around Lake Malawi and the Shire valley. Trade and marriage alliances connected lakeshore and inland populations.
Source confidence: high