The genomic history of southeastern Europe

Mathieson I; Alpaslan-Roodenberg S; Posth C; Szécsényi-Nagy A; Rohland N; Mallick S; Olalde I; Broomandkhoshbacht N; Candilio F; Cheronet O; Fernandes D; Ferry M; Gamarra B; Fortes GG; Haak W; Harney E; Jones E; Keating D; Krause-Kyora B; Kucukkalipci I; Michel M; Mittnik A; Nägele K; Novak M; Oppenheimer J; Patterson N; Pfrengle S; Sirak K; Stewardson K; Vai S; Alexandrov S; Alt KW; Andreescu R; Antonović D; Ash A; Atanassova N; Bacvarov K; Gusztáv MB; Bocherens H; Bolus M; Boroneanţ A; Boyadzhiev Y; Budnik A; Burmaz J; Chohadzhiev S; Conard NJ; Cottiaux R; Čuka M; Cupillard C; Drucker DG; Elenski N; Francken M; Galabova B; Ganetsovski G; Gély B; Hajdu T; Handzhyiska V; Harvati K; Higham T; Iliev S; Janković I; Karavanić I; Kennett DJ; Komšo D; Kozak A; Labuda D; Lari M; Lazar C; Leppek M; Leshtakov K; Vetro DL; Los D; Lozanov I; Malina M; Martini F; McSweeney K; Meller H; Menđušić M; Mirea P; Moiseyev V; Petrova V; Price TD; Simalcsik A; Sineo L; Šlaus M; Slavchev V; Stanev P; Starović A; Szeniczey T; Talamo S; Teschler-Nicola M; Thevenet C; Valchev I; Valentin F; Vasilyev S; Veljanovska F; Venelinova S; Veselovskaya E; Viola B; Virag C; Zaninović J; Zäuner S; Stockhammer PW; Catalano G; Krauß R; Caramelli D; Zariņa G; Gaydarska B; Lillie M; Nikitin AG; Potekhina I; Papathanasiou A; Borić D; Bonsall C; Krause J; Pinhasi R; Reich D

doi:10.1038/nature25778

Research Publication

The genomic history of southeastern Europe

Mathieson I, Alpaslan-Roodenberg S, Posth C et al.

29466330 PubMed ID

117 Authors

03/08/2018 Published

282 Samples

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Chapter I

Publication Details

Comprehensive information about this research publication

Authors 117 researchers

Publication Date 03/08/2018

Geographic Region Europe

URL https://pubmed.ncbi.nlm.nih.gov/29466330/

Authors

MI

Mathieson I

AS

Alpaslan-Roodenberg S

PC

Posth C

SA

Szécsényi-Nagy A

RN

Rohland N

MS

Mallick S

OI

Olalde I

BN

Broomandkhoshbacht N

CF

Candilio F

CO

Cheronet O

FD

Fernandes D

FM

Ferry M

GB

Gamarra B

FG

Fortes GG

HW

Haak W

HE

Harney E

JE

Jones E

KD

Keating D

KB

Krause-Kyora B

KI

Kucukkalipci I

MM

Michel M

MA

Mittnik A

NK

Nägele K

NM

Novak M

OJ

Oppenheimer J

PN

Patterson N

PS

Pfrengle S

SK

Sirak K

SK

Stewardson K

VS

Vai S

AS

Alexandrov S

AK

Alt KW

AR

Andreescu R

AD

Antonović D

AA

Ash A

AN

Atanassova N

BK

Bacvarov K

GM

Gusztáv MB

BH

Bocherens H

BM

Bolus M

BA

Boroneanţ A

BY

Boyadzhiev Y

BA

Budnik A

BJ

Burmaz J

CS

Chohadzhiev S

CN

Conard NJ

CR

Cottiaux R

ČM

Čuka M

CC

Cupillard C

DD

Drucker DG

EN

Elenski N

FM

Francken M

GB

Galabova B

GG

Ganetsovski G

GB

Gély B

HT

Hajdu T

HV

Handzhyiska V

HK

Harvati K

HT

Higham T

IS

Iliev S

JI

Janković I

KI

Karavanić I

KD

Kennett DJ

KD

Komšo D

KA

Kozak A

LD

Labuda D

LM

Lari M

LC

Lazar C

LM

Leppek M

LK

Leshtakov K

VD

Vetro DL

LD

Los D

LI

Lozanov I

MM

Malina M

MF

Martini F

MK

McSweeney K

MH

Meller H

MM

Menđušić M

MP

Mirea P

MV

Moiseyev V

PV

Petrova V

PT

Price TD

SA

Simalcsik A

SL

Sineo L

ŠM

Šlaus M

SV

Slavchev V

SP

Stanev P

SA

Starović A

ST

Szeniczey T

TS

Talamo S

TM

Teschler-Nicola M

TC

Thevenet C

VI

Valchev I

VF

Valentin F

VS

Vasilyev S

VF

Veljanovska F

VS

Venelinova S

VE

Veselovskaya E

VB

Viola B

VC

Virag C

ZJ

Zaninović J

ZS

Zäuner S

SP

Stockhammer PW

CG

Catalano G

KR

Krauß R

CD

Caramelli D

ZG

Zariņa G

GB

Gaydarska B

LM

Lillie M

NA

Nikitin AG

PI

Potekhina I

PA

Papathanasiou A

BD

Borić D

BC

Bonsall C

KJ

Krause J

PR

Pinhasi R

RD

Reich D

View on PubMed

Chapter II

Abstract

Summary of the research findings

Farming was first introduced to Europe in the mid-seventh millennium bc, and was associated with migrants from Anatolia who settled in the southeast before spreading throughout Europe. Here, to understand the dynamics of this process, we analysed genome-wide ancient DNA data from 225 individuals who lived in southeastern Europe and surrounding regions between 12000 and 500 bc. We document a west-east cline of ancestry in indigenous hunter-gatherers and, in eastern Europe, the early stages in the formation of Bronze Age steppe ancestry. We show that the first farmers of northern and western Europe dispersed through southeastern Europe with limited hunter-gatherer admixture, but that some early groups in the southeast mixed extensively with hunter-gatherers without the sex-biased admixture that prevailed later in the north and west. We also show that southeastern Europe continued to be a nexus between east and west after the arrival of farmers, with intermittent genetic contact with steppe populations occurring up to 2,000 years earlier than the migrations from the steppe that ultimately replaced much of the population of northern Europe.

Chapter III

Ancient DNA Samples

282 ancient DNA samples referenced in this publication

282 Samples

Sample ID	Date/Era	Country	Locality	Sex	mtDNA	Y-DNA
I5070	5209 BCE	Austria	Asparn Schletz	M	`K1a1a*`	`C-FT251331`
I5204	5500 BCE	Austria	Asparn Schletz	M	`J1c2`	`G-Z39358`
I5205	5100 BCE	Austria	Asparn Schletz	F	`H`
I5206	5100 BCE	Austria	Asparn Schletz	F	`T2b`
I5207	5100 BCE	Austria	Asparn Schletz	M	`H67`	`J-Z6048`
I5068	5500 BCE	Austria	Kleinhadersdorf Flur Marchleiten	M	`T2b23`	`J-Z6048`
I5069	5205 BCE	Austria	Kleinhadersdorf Flur Marchleiten	F	`N1a1a1*`
I5208	5500 BCE	Austria	Asparn Schletz	F	`K1b1a`
I5207	5100 BCE	Austria	Asparn Schletz	M	`H67`	`J-Z6048`
I5070	5209 BCE	Austria	Asparn Schletz	M	`K1a1a*`
I5204	5500 BCE	Austria	Asparn Schletz	M	`J1c2`
I5205	5100 BCE	Austria	Asparn Schletz	F	`H`
I5206	5100 BCE	Austria	Asparn Schletz	F	`T2b`
I2176	3340 BCE	Bulgaria	Smyadovo	M	`U1a1`	`I-Y87044`
I2181	4606 BCE	Bulgaria	Smyadovo	M	`HV15`	`R-M269`
I2215	5800 BCE	Bulgaria	Malak Preslavets	M	`T2b`	`R-M459`
I2216	5800 BCE	Bulgaria	Malak Preslavets	F	`J2b1`
I2426	4446 BCE	Bulgaria	Sushina	M	`K1`	`F-M89`
I0704	6059 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	F	`T2b`
I1296	5800 BCE	Bulgaria	Malak Preslavets	M	`U5a2`	`C-BY1463`
I0698	6000 BCE	Bulgaria	Yabalkovo. Dimitrovgrad. Haskovo	M	`H`	`G-Z6488`
I2529	5723 BCE	Bulgaria	Yabalkovo. Dimitrovgrad. Haskovo	M	`T1a`	`I-M223`
I1298	6057 BCE	Bulgaria	Ohoden	F	`K1-a`
I2431	4784 BCE	Bulgaria	Ivanovo	M	`N1b2`	`G-S10654`
I2526	5468 BCE	Bulgaria	Samovodene	F	`T2e`
I2423	4536 BCE	Bulgaria	Smyadovo	F	`H`
I2424	4445 BCE	Bulgaria	Smyadovo	F	`U4a`
I2430	4602 BCE	Bulgaria	Smyadovo	M	`K1a26`	`R-FGC21056`
I2181	4606 BCE	Bulgaria	Smyadovo	M	`HV15`	`R-M269`
I2426	4446 BCE	Bulgaria	Sushina	M	`K1`	`F-M89`
I2425	4676 BCE	Bulgaria	Sushina	F	`T2c1b`
I2427	4446 BCE	Bulgaria	Sushina	F	`H1j`
I2509	4449 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	F	`K1a2e`
I2519	4341 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	F	`U5b2a1a`
I0785	4451 BCE	Bulgaria	Yunatsite	F	`H7`
I0781	4537 BCE	Bulgaria	Yunatsite	F	`K1-a`
ANI152	4589 BCE	Bulgaria	Varna	M	`U2`	`T-M184`
ANI153	4550 BCE	Bulgaria	Varna	M	`U4`	`R-V88`
ANI160	4684 BCE	Bulgaria	Varna	M	`H1ag`	`G-L30`
ANI163	4687 BCE	Bulgaria	Varna	F	`H7a1`
merge_ANI159_ANI181	4714 BCE	Bulgaria	Varna	M	`T2b2b`	`G-PF3359`
Bul6	3400 BCE	Bulgaria	Nova Zagora. Beli Breyag	M		`I-S12991`
Bul8	3400 BCE	Bulgaria	Nova Zagora. Beli Breyag	M		`I-P215`
I2165	3072 BCE	Bulgaria	Merichleri. Kairyaka necropolis	M	`T2f`	`I-S12195`
Bul4	3011 BCE	Bulgaria	Nova Zagora. Mednikarovo	M		`I-L703`
I2175	3335 BCE	Bulgaria	Smyadovo	M	`K1c1`	`I-Y87044`
I2176	3340 BCE	Bulgaria	Smyadovo	M	`U1a1`	`I-Y87044`
I2520	3331 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	M	`H*`	`H-FT369003`
I2510	2907 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	M	`H4a1*`	`G-FTA37257`
Bul10	3092 BCE	Bulgaria	Sabrano (Sliven province, Nova Zagora municipality)	F
I12835	3092 BCE	Bulgaria	Nova Zagora. Sabrano	F
I5769	771 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	F	`H-d9`
I2163	1866 BCE	Bulgaria	Merichleri. Kairyaka necropolis	M	`U5a2`	`R-Z93`
I1298	6057 BCE	Bulgaria	Ohoden	F	`K1-a`
I0704	6059 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	F	`T2b`
I2521	5621 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	M	`H*`	`G-F2214`
I0706	6025 BCE	Bulgaria	Veliko Tarnovo. Dzhulyunitsa	M	`K1a4b`	`C-PH428`
I0698	6000 BCE	Bulgaria	Yabalkovo. Dimitrovgrad. Haskovo	M	`H`	`G-Z6488`
I2529	5723 BCE	Bulgaria	Yabalkovo. Dimitrovgrad. Haskovo	M	`T1a`	`I-M223`
I0679	5723 BCE	Bulgaria	Krepost	F	`H5`
I1108	5800 BCE	Bulgaria	Malak Preslavets	M	`T2e`	`T-L454`
I1109	5800 BCE	Bulgaria	Malak Preslavets	F	`J2b1`
I1113	5800 BCE	Bulgaria	Malak Preslavets	F	`U5a1c`
I1295	5800 BCE	Bulgaria	Malak Preslavets	M	`J1c`	`G-P303`
I1297	5800 BCE	Bulgaria	Malak Preslavets	F	`H5b`
I0700	5800 BCE	Bulgaria	Malak Preslavets	M	`T2e`	`T-L454`
I3879	5800 BCE	Bulgaria	Malak Preslavets	M	`H`	`G-P303`
I2215	5800 BCE	Bulgaria	Malak Preslavets	M	`T2b`	`R-M459`
I2216	5800 BCE	Bulgaria	Malak Preslavets	F	`J2b1`
I1296	5800 BCE	Bulgaria	Malak Preslavets	M	`U5a2`	`C-BY1463`
I3313	805 BCE	Croatia	Jazinka Cave	F	`HV0e*`
I5071	5708 BCE	Croatia	Kargadur	F	`H5a`
I5072	5657 BCE	Croatia	Kargadur	M	`H7c`	`G-PF3177`
I3499	2885 BCE	Croatia	Beli Manastir-Popova zemlja	M	`T2e`	`R-M12149`
I4175	3000 BCE	Croatia	Vučedol Tell	F	`U4a`
I1875	7321 BCE	Croatia	Vela Spila	F	`U5b2b`
I4331	1618 BCE	Croatia	Veliki Vanik	M	`I1a1-a*`	`J-Z38240`
I4332	1613 BCE	Croatia	Veliki Vanik	F	`W3a1`
I4168	4777 BCE	Croatia	Beli Manastir-Popova zemlja	F	`N1a1a1`
I3498	5834 BCE	Croatia	Beli Manastir-Popova zemlja	M	`U8b1b1`	`C-PH428`
I5077	5208 BCE	Croatia	Osijek. Hermanov Vinograd	M	`U5a1c1*`	`G-FGC55322`
I5078	4699 BCE	Croatia	Osijek. Hermanov Vinograd	M	`H10-a*`	`J-Z36829`
I3433	5975 BCE	Croatia	Zemunica Cave	F	`H1`
I3947	5985 BCE	Croatia	Zemunica Cave	M	`K1b1a`	`C-F1628`
I3948	6007 BCE	Croatia	Zemunica Cave	M	`N1a1`	`E-L618`
I4167	4792 BCE	Croatia	Beli Manastir-Popova zemlja	M	`U5b2b`	`I-FT355000`
I5079	3651 BCE	Croatia	Radovanci	F	`J1c2aa`
I2792	2872 BCE	Croatia	Vučedol Tell	M	`T2c2`	`G-Z6488`
Iboussieres25-1	10050 BCE	France	Aven des Iboussières à Malataverne. Rhône-Alpes	M	`U5b2a`	`I-P215`
Iboussieres31-2	10050 BCE	France	Aven des Iboussières à Malataverne. Rhône-Alpes	M	`U5b1`	`R-M173`
BerryAuBac	5368 BCE	France	Berry Au Bac	M	`U5b1a`	`I-M170`
BerryAuBac	5368 BCE	France	Berry Au Bac	M	`U5b1a`	`I-M170`
Iboussieres25-1	10050 BCE	France	Aven des Iboussières à Malataverne. Rhône-Alpes	M	`U5b2a`	`I-P215`
Iboussieres31-2	10050 BCE	France	Aven des Iboussières à Malataverne. Rhône-Alpes	M	`U5b1`	`R-M173`
Falkenstein	7472 BCE	Germany	Swabian Jura. Baden-Wurttemberg. Falkenstein-Höhle	M	`U5a2c`	`I-CTS616`
Falkenstein	7472 BCE	Germany	Swabian Jura. Baden-Wurttemberg. Falkenstein-Höhle	M	`U5a2c`	`I-CTS616`
I5427	6009 BCE	Greece	Diros. Alepotrypa Cave	F	`K1a24`
I2318	4049 BCE	Greece	Argolid. Franchthi Cave	F	`H2`
I3920	3938 BCE	Greece	Diros. Alepotrypa Cave	F	`H`
I3708	5500 BCE	Greece	Diros. Alepotrypa Cave	F	`T1a`
I3709	4036 BCE	Greece	Diros. Alepotrypa Cave	F	`K1b1a`
I2158	12350 BCE	Italy	Sicily. Egadi Islands. Favignana Island. Grotta d'Oriente	F	`U2'3'4'7'8'9`
I4628	5304 BCE	Latvia	Zvejnieki	M	`U5a2d4`	`R-FTA35786`
I4432	6072 BCE	Latvia	Zvejnieki	M	`U5a2c*`	`R-A11291`
I4434	5611 BCE	Latvia	Zvejnieki	M	`U5a2d3`	`R-FTA35785`
I4438	5375 BCE	Latvia	Zvejnieki	M	`U4b1a2*`	`I-FTA37476`
I4439	5769 BCE	Latvia	Zvejnieki	M	`U5b1d1*`	`R-FTA35787`
I4440	5216 BCE	Latvia	Zvejnieki	M	`U4a1o`	`I-FTA37476`
I4441	4841 BCE	Latvia	Zvejnieki	M	`U5a1c3b`	`I-L801`
I4550	6633 BCE	Latvia	Zvejnieki	M	`U5b2a7`	`Q-Y2700`
I4551	5788 BCE	Latvia	Zvejnieki	M	`U5a2f1`	`I-FT370114`
I4552	5477 BCE	Latvia	Zvejnieki	F	`U5a1d*`
I4553	5480 BCE	Latvia	Zvejnieki	M	`U5a1c`	`I-FT370114`
I4595	5729 BCE	Latvia	Zvejnieki	F	`U2e1`
I4596	6069 BCE	Latvia	Zvejnieki	M	`U5a2d3`	`I-PH5234`
I4630	7471 BCE	Latvia	Zvejnieki	M	`U5a2c`	`R-P297`
I4632	6465 BCE	Latvia	Zvejnieki	F	`U5a1c3b*`
I4629	3091 BCE	Latvia	Zvejnieki	F	`U5a1b`
I4626	5838 BCE	Latvia	Zvejnieki	M	`U2e1`	`R-A11291`
I4435	4214 BCE	Latvia	Zvejnieki	F	`U4a1p`
I4436	4318 BCE	Latvia	Zvejnieki	M	`U4a1p`	`R-FTA35785`
I4437	4330 BCE	Latvia	Zvejnieki	F	`U5a1d2*`
I4554	4048 BCE	Latvia	Zvejnieki	F	`U4d1`
I4627	4311 BCE	Latvia	Zvejnieki	M	`U4a1`	`R-M73`
I0001	6221 BCE	Luxembourg	Echternach	M	`U5b1a`	`I-V4921`
I0676	5979 BCE	North Macedonia	Skopje. Sopište. Govrlevo	M	`J1c1`	`G-PF3430`
I2405	3337 BCE	Poland	Kierzkowo	M	`W5`	`I-M223`
I2435	3100 BCE	Poland	Kierzkowo	M	`H28`	`I-CTS2257`
I2407	3096 BCE	Poland	Kierzkowo	M	`H28a`	`I-CTS2257`
I2405	3337 BCE	Poland	Kierzkowo	M	`W5`	`I-M223`
I2435	3100 BCE	Poland	Kierzkowo	M	`H28`	`I-CTS2257`
I2407	3096 BCE	Poland	Kierzkowo	M	`H28a`	`I-CTS2257`
I2433	3100 BCE	Poland	Kierzkowo	F	`H28`
I2434	3400 BCE	Poland	Kierzkowo	F	`U5b1d1`
I2440	3100 BCE	Poland	Kierzkowo	M	`H1b`	`I-CTS2257`
I2441	3400 BCE	Poland	Kierzkowo	M	`K1b1a1`	`I-L801`
I2403	2868 BCE	Poland	Kierzkowo	M	`U5b2b1`	`I-Z161`
I4655	7059 BCE	Romania	Iron Gates. Schela Cladovei	M	`K1`	`R-M173`
I4081	7581 BCE	Romania	Iron Gates. Ostrovul Corbului	M	`H13`	`R-L754`
I2533	5480 BCE	Romania	Carcea	F	`J1c5*`
I2532	5721 BCE	Romania	Cotatcu	M	`K1a2`	`G-CTS8176`
I4582	7026 BCE	Romania	Iron Gates. Ostrovul Corbului	F	`K1i`
I4081	7581 BCE	Romania	Iron Gates. Ostrovul Corbului	M	`H13`	`R-L754`
I4655	7059 BCE	Romania	Iron Gates. Schela Cladovei	M	`K1`	`R-M173`
I4607	7446 BCE	Romania	Iron Gates. Schela Cladovei	M	`U5a2`	`I-S2599`
I5436	7451 BCE	Romania	Iron Gates. Schela Cladovei	F
I5411	7000 BCE	Romania	Iron Gates. Schela Cladovei	M	`U5a1c1`	`R-FT360000`
I2534	5810 BCE	Romania	Magura Buduiasca. Teleor 3	F	`K1`
I5408	7022 BCE	Romania	Iron Gates. Ostrovul Corbului	M	`K1i`	`R-FT360000`
I4088	4332 BCE	Romania	Urziceni	F	`K2a`
I4089	3765 BCE	Romania	Urziceni	M	`J1c*`	`C-Z29332`
I5409	9300 BCE	Serbia	Padina	F	`U5a1c`
I4881	6570 BCE	Serbia	Vlasac	M	`U4b1b1g1`	`I-L801`
I4914	6360 BCE	Serbia	Hajduka Vodenica	M	`U5a1c1*`	`I-FTA37531`
I4657	9755 BCE	Serbia	Vlasac	F	`K1c`
I4665	6205 BCE	Serbia	Lepenski Vir	F	`J2b1`
I4666	6222 BCE	Serbia	Lepenski Vir	M	`H40`	`R-PF6323`
I5773	8240 BCE	Serbia	Vlasac	M	`U4a`	`I-M170`
I0633	5610 BCE	Serbia	Vojvodina. Hrtkovci. Gomolova	M	`HV`	`G-FGC34625`
I0634	4712 BCE	Serbia	Vojvodina. Hrtkovci. Gomolova	M	`K1a4`	`G-FGC34625`
I0634	4712 BCE	Serbia	Vojvodina. Hrtkovci. Gomolova	M	`K1a4`	`G-FGC34625`
I0633	5610 BCE	Serbia	Vojvodina. Hrtkovci. Gomolova	M	`HV`	`G-FGC34625`
I1131	4653 BCE	Serbia	Vojvodina. Hrtkovci. Gomolova	M	`H*`	`G-FT370002`
I4918	6000 BCE	Serbia	Saraorci-Jezava	F	`K1a4a1*`
I4915	6340 BCE	Serbia	Hajduka Vodenica	M	`U5b2b-a3a`	`I-S2599`
I4916	7035 BCE	Serbia	Hajduka Vodenica	M	`U5b2b-a3a`	`R-FT360000`
I4917	6220 BCE	Serbia	Hajduka Vodenica	F	`U5a1c3b`
I5401	7076 BCE	Serbia	Hajduka Vodenica	M	`U5a1`	`I-S2599`
I5402	6361 BCE	Serbia	Hajduka Vodenica	M	`U5a1c1`	`I-CTS4348`
I4914	6360 BCE	Serbia	Hajduka Vodenica	M	`U5a1c1*`	`I-FTA37531`
I5407	6300 BCE	Serbia	Lepenski Vir	F	`H13*`
I5232	6061 BCE	Serbia	Padina	M	`K1-a4*`	`R-FTA35718`
I5233	6224 BCE	Serbia	Padina	F	`U5b1d1*`
I5234	9300 BCE	Serbia	Padina	F	`U4a7`
I5235	9221 BCE	Serbia	Padina	M	`U5b2c*`	`R-FTA35749`
I5236	8291 BCE	Serbia	Padina	M	`U5a2d*`	`I-BY33164`
I5237	9300 BCE	Serbia	Padina	M	`U5a2f1`	`R-FT370000`
I5238	8265 BCE	Serbia	Padina	F	`K1g1`
I5239	8535 BCE	Serbia	Padina	F	`U5a2d*`
I5240	9140 BCE	Serbia	Padina	M	`U5a1c3*`	`R-FTA35749`
I5241	9302 BCE	Serbia	Padina	F	`U5a2a`
I5242	8805 BCE	Serbia	Padina	F	`U5a1m*`
I5244	9115 BCE	Serbia	Padina	F	`K1*`
I5409	9300 BCE	Serbia	Padina	F	`U5a1c`
I5773	8240 BCE	Serbia	Vlasac	M	`U4a`	`I-M170`
I4881	6570 BCE	Serbia	Vlasac	M	`U4b1b1g1`	`I-L801`
I4660	7940 BCE	Serbia	Vlasac	F	`U8b1b`
I4870	7045 BCE	Serbia	Vlasac	M	`K1c`	`I-BY32198`
I4657	9755 BCE	Serbia	Vlasac	F	`K1c`
I4871	7100 BCE	Serbia	Vlasac	F	`U5b2a1a`
I4872	7100 BCE	Serbia	Vlasac	F	`U5a1c`
I4873	6012 BCE	Serbia	Vlasac	F	`U5a2-a*`
I4874	6640 BCE	Serbia	Vlasac	F	`U5b2a7`
I4875	6615 BCE	Serbia	Vlasac	F	`U5b1d1*`
I4876	6635 BCE	Serbia	Vlasac	F	`U5a2d*`
I4877	6645 BCE	Serbia	Vlasac	F	`U5b1d1*`
I4878	5995 BCE	Serbia	Vlasac	M	`U4a5*`	`I-Y7240`
I4880	6000 BCE	Serbia	Vlasac	M	`U4b1b1g*`	`I-FTA37515`
I4882	6200 BCE	Serbia	Vlasac	M	`U4b1b1g*`	`I-FTA37515`
I5771	6500 BCE	Serbia	Vlasac	M	`U5a1c1`	`I-P37`
I5772	7100 BCE	Serbia	Vlasac	M	`U5a2a`	`R-V88`
I5405	5836 BCE	Serbia	Lepenski Vir	F	`HV`
I4665	6205 BCE	Serbia	Lepenski Vir	F	`J2b1`
I4666	6222 BCE	Serbia	Lepenski Vir	M	`H40`	`R-PF6323`
I2111	3765 BCE	Ukraine	Verteba Cave	M	`HV`	`G-P15`
I3141	3300 BCE	Ukraine	Shevchenko. OAE-2003	F	`H15b1`
I3151	3950 BCE	Ukraine	Verteba Cave	M	`U8b1b`	`E-M78`
I4112	5500 BCE	Ukraine	Dereivka I	M	`U5a2a`	`R-PF6287`
I3712	5512 BCE	Ukraine	Volniensky. Vilnianka	M	`U4b1a`	`I-CTS10057`
I3713	5201 BCE	Ukraine	Volniensky. Vilnianka	M	`U4b1a`	`I-P215`
I3716	5473 BCE	Ukraine	Volniensky. Vilnianka	M	`U5b2a1a`	`I-M223`
I5868	5471 BCE	Ukraine	Volniensky. Vilnianka	M	`U4d`	`I-CTS616`
I5872	5481 BCE	Ukraine	Volniensky. Vilnianka	M	`U5a2a`	`I-L703`
I5873	5472 BCE	Ukraine	Volniensky. Vilnianka	F	`U4b`
I5878	5301 BCE	Ukraine	Dereivka I	M	`U4b1a1a`	`R-BY17643`
I5881	5215 BCE	Ukraine	Dereivka I	M	`U5a1b`	`R-V88`
I5885	6392 BCE	Ukraine	Dereivka I	F	`U5b2b`
I5886	5313 BCE	Ukraine	Dereivka I	M	`U4a`	`I-L702`
I5888	5500 BCE	Ukraine	Dereivka I	M	`U5b2b1`	`I-P214`
I5889	5314 BCE	Ukraine	Dereivka I	F	`U5a2a`
I5891	5467 BCE	Ukraine	Dereivka I	M	`U4d`	`R-V88`
I5893	5371 BCE	Ukraine	Dereivka I	M	`U5a2a`	`R-PF6287`
I5957	5470 BCE	Ukraine	Volniensky. Vilnianka	M	`T2`	`I-P222`
I6133	6500 BCE	Ukraine	Volniensky. Vilnianka	F	`U5b2a1a`
I3719	4983 BCE	Ukraine	Dereivka I	M	`H1`	`I-P222`
I5884	2886 BCE	Ukraine	Dereivka I	M	`U5a2b`	`R-M12149`
I1737	8543 BCE	Ukraine	Vasil'evka	F	`U5a2`
ILK002	2889 BCE	Ukraine	Ilatka	M	`T2b`	`I-Y167771`
I5884	2886 BCE	Ukraine	Dereivka I	M	`U5a2b`	`R-M12149`
I1917	3096 BCE	Ukraine	Ozera. OAE-99	F	`R0a1`
I2105	3300 BCE	Ukraine	Shevchenko. OAE-2003	F	`T1a1*`
I3141	3300 BCE	Ukraine	Shevchenko. OAE-2003	F	`H15b1`
I3151	3950 BCE	Ukraine	Verteba Cave	M	`U8b1b`	`E-M78`
I2111	3765 BCE	Ukraine	Verteba Cave	M	`HV`	`G-P15`
I2110	3911 BCE	Ukraine	Verteba Cave	M	`T2b`
I1926	3769 BCE	Ukraine	Verteba Cave	M	`H5a`	`G-P303`
I4110	3509 BCE	Ukraine	Dereivka I	F	`J2b1`
I5882	3308 BCE	Ukraine	Dereivka I	F	`U5a2a`
ILK003	2901 BCE	Ukraine	Ilatka	F	`J1c`
ILK001	2900 BCE	Ukraine	Ilatka	M	`J1c3`	`I-Y167771`
I5876	7040 BCE	Ukraine	Dereivka I	M	`U5a2a`	`R-YP5056`
I5885	6392 BCE	Ukraine	Dereivka I	F	`U5b2b`
I1737	8543 BCE	Ukraine	Vasil'evka	F	`U5a2`
I1733	9000 BCE	Ukraine	Vasil'evka	F	`U4b`
I1763	8282 BCE	Ukraine	Vasil'evka	M	`U5b2d`	`I-FT344596`
I1819	9107 BCE	Ukraine	Vasil'evka	M	`U5b2e*`	`R-M459`
I6561	2134 BCE	Ukraine	Alexandria	M	`H2a1a`	`R-F2597`
I3718	5359 BCE	Ukraine	Dereivka I	M	`U5a1b`	`R-BY17644`
I5879	5324 BCE	Ukraine	Dereivka I	M	`U5a1b`	`R-BY17643`
I5883	5209 BCE	Ukraine	Dereivka I	M	`U4a`	`R-BY17643`
I5890	5291 BCE	Ukraine	Dereivka I	M	`U5a1b`	`R-BY17643`
I5892	5301 BCE	Ukraine	Dereivka I	M	`U4a1`	`R-L754`
I5875	5295 BCE	Ukraine	Dereivka I	M	`U4a1`	`I-L703`
I4111	4725 BCE	Ukraine	Dereivka I	F	`U4d`
I4114	5474 BCE	Ukraine	Dereivka I	M	`U5a1`	`R-BY17643`
I3717	5371 BCE	Ukraine	Dereivka I	M	`U5a2a`	`I-L703`
I5878	5301 BCE	Ukraine	Dereivka I	M	`U4b1a1a`	`R-BY17643`
I3719	4983 BCE	Ukraine	Dereivka I	M	`H1`	`I-P222`
I5888	5500 BCE	Ukraine	Dereivka I	M	`U5b2b1`	`I-P214`
I5881	5215 BCE	Ukraine	Dereivka I	M	`U5a1b`	`R-V88`
I5893	5371 BCE	Ukraine	Dereivka I	M	`U5a2a`	`R-PF6287`
I5886	5313 BCE	Ukraine	Dereivka I	M	`U4a`	`I-L702`
I5889	5314 BCE	Ukraine	Dereivka I	F	`U5a2a`
I5891	5467 BCE	Ukraine	Dereivka I	M	`U4d`	`R-V88`
I4112	5500 BCE	Ukraine	Dereivka I	M	`U5a2a`	`R-PF6287`
I1736	6242 BCE	Ukraine	Vasil'evka	F	`U5b2b1`
I1734	6300 BCE	Ukraine	Vasil'evka	M	`U5b2d`	`R-PF6362`
I3714	5500 BCE	Ukraine	Volniensky. Vilnianka	M	`U4d`	`I-P222`
I3715	5636 BCE	Ukraine	Volniensky. Vilnianka	M	`U2e1`	`I-S22311`
I5869	5612 BCE	Ukraine	Volniensky. Vilnianka	M	`U4b1b1`	`I-P222`
I5870	5475 BCE	Ukraine	Volniensky. Vilnianka	M	`U4b1b1`	`I-S12195`
I5873	5472 BCE	Ukraine	Volniensky. Vilnianka	F	`U4b`
I5868	5471 BCE	Ukraine	Volniensky. Vilnianka	M	`U4d`	`I-CTS616`
I6133	6500 BCE	Ukraine	Volniensky. Vilnianka	F	`U5b2a1a`
I3712	5512 BCE	Ukraine	Volniensky. Vilnianka	M	`U4b1a`	`I-CTS10057`
I3713	5201 BCE	Ukraine	Volniensky. Vilnianka	M	`U4b1a`	`I-P215`
I5957	5470 BCE	Ukraine	Volniensky. Vilnianka	M	`T2`	`I-P222`
I5872	5481 BCE	Ukraine	Volniensky. Vilnianka	M	`U5a2a`	`I-L703`
I3716	5473 BCE	Ukraine	Volniensky. Vilnianka	M	`U5b2a1a`	`I-M223`
I1738	5475 BCE	Ukraine	Vovnigi	M	`U5a2`	`I-L703`
I1732	5372 BCE	Ukraine	Vovnigi	F	`U4*`
I1378	5400 BCE	Ukraine	Vovnigi	M	`U4b`	`I-L702`

Chapter IV

Analysis

Comprehensive review of ancestry and genetic findings

Important Disclaimer: This review has been performed semi-automatically and is provided for informational purposes only. While we strive for accuracy, this analysis may contain errors, omissions, or misinterpretations of the original research. DNA Genics disclaims all liability for any inaccuracies, errors, or consequences arising from the use of this information. Users should independently verify all information and consult original research publications before making any decisions based on this content. This analysis is not intended as a substitute for professional scientific review or medical advice.

The genomic history of southeastern Europe

Publication Details

Authors

Abstract

Ancient DNA Samples

Analysis

Analysis In Progress

Summary

Key Findings

Ancestry Insights

Traits Analysis

Historical Context

Scientific Assessment

Explore More Research

The genomic history of southeastern Europe

Publication Details

Authors

Abstract

Ancient DNA Samples

Analysis

Analysis In Progress

Summary

Key Findings

Ancestry Insights

Traits Analysis

Historical Context

Scientific Assessment

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